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This observed natural heterogeneity is significantly larger than the modelled heterogeneity, in particular in the case of the evapotranspiration rate. Concerning the inter-annual variability, the correlation between modelled and observed yearly anomalies with the mean GPP and ET at the different sites over the time series provided in Table 1 is weak. The correlation coefficient r is equal to 0.62 (p?[http://en.wikipedia.org/wiki/Ribosomal_s6_kinase S6 Kinase] the observed and modelled seasonal variability. In Fig. 3(a) and Table 4 the partitioning of the modelled yearly evapotranspiration [http://www.selleckchem.com/products/gsk-j4-hcl.html GSK J4 molecular weight] flux between its different components is presented. Results show that the contribution of the transpiration, soil and intercepted water evaporation to the total evapotranspiration rate is not in line with what can be found in the literature. According to a global process-based study performed by Miralles et al. (2011), the transpiration accounts for 80% of the annual land evaporation, interception loss for 11%, bare soil evaporation for 7% and snow sublimation for 2%. More specifically for Europe, the interception loss represents 18%. A study by Choudhury and DiGirolamo [http://www.selleckchem.com/products/MLN-2238.html this website] (1998), based on 132 geographically distributed catchments, estimated the ratio of transpiration to evapotranspiration around 0.65�C0.7 for forest and 0.6 for grassland. At the selected sites in this study, the transpiration accounts on average for only 42%, the soil evaporation for 34% and the intercepted water evaporation for 24% (see Table 4). Very few measurements have been performed at site specific locations. Wilson et al. (2001) measured the contribution of transpiration, intercepted water and soil evaporation in temperate deciduous forests and found it to be 60%, 20% and 20%, respectively. A similar contribution of soil evaporation was observed for the temperate broad-leaved forest in Hainich (Bittner et al., 2010). During the summer however, the contribution of the soil dropped to 10�C13%. At Brasschaat, the observed soil evaporation contributes about 30% to the total evapotranspiration (Meiresonne et al., 2003). At Vielsalm in 2011, the transpiration represented 53% and the intercepted water evaporation 13% (personal communication with R. Soubie and C. Vincke �C 12/04/2013). Averaged over 14 stands in the North of Belgium, the contribution of transpiration is around 65%, that of intercepted water evaporation around 25% and of soil water evaporation 10% (Verstraeten et al., 2005). Sutanto et al. (2012) measured the evaporation fluxes during simulated summer conditions on a grassland. The fraction of transpiration, soil and intercepted water evaporation were 77.7%, 12.2% and 10.