N Fig. 15B); tarsal claw comparatively huge and stout having a

The genus Asthenopus has been distinguished by means with the following characters (Dom guez 1988): 1) ratio foreleg/FW male: 3/5?/5; two) male foretarsus two.5 occasions longer than foretibia; 3) foretarsal segment 2 equivalent to the other folks, and 2/3 the length of tibia); four) ratio length of Rs stem/fork to margin 1/4 (or fork Rs at 2/10 from base to margin) ; 5) cubital intercalaries slightly diverging toward hind margin, ICu2 and ICu1 basally fused to CuA by cross veins; 6) ICu2 ending at anal margin or within the tornus, 7) marginal intercalary veins absent; 8) forceps ratio width/length: 1/7; 8) penes Win 63843 clinical trials robust on basal 2/3; 9) MA fork base to margin 10/100; 11) IMP P1 not fused basally; 12) MP2-IMP equivalent in length, not fused; 13) foretarsal claws of male not so expanded distally (as in Asthenopodes). Coxae I and II directed ventrally, coxae III directed laterally. Abdomen. Gill I lowered in size, double, each portions subequal in length and width. Gills II II properly developed, ventral portion smaller than dorsal portion; tergum X with effectively developed ventral spine on posterior margin (not visible dorsally, Fig. 15E). Caudal filaments brief (curved in mature nymphs) with whorls of stout spines and basic setae at joinings. Distribution. Amazonas and Parana biogeographic subregions (Argentina, Bolivia, Brazil, Colombia, Ecuador, Peru). Discussion. The genus Asthenopus has been distinguished by implies with the following characters (Dom guez 1988): 1) ratio foreleg/FW male: 3/5?/5; 2) male foretarsus 2.5 occasions longer than foretibia; three) foretarsal segment 2 equivalent to the other individuals, and 2/3 the length of tibia); 4) ratio length of Rs stem/fork to margin 1/4 (or fork Rs at 2/10 from base to margin) ; 5) cubital intercalaries slightly diverging toward hind margin, ICu2 and ICu1 basally fused to CuA by cross veins; six) ICu2 ending at anal margin or inside the tornus, 7) marginal intercalary veins absent; eight) forceps ratio width/length: 1/7; 8) penes robust on basal 2/3; 9) MA fork base to margin 10/100; 11) IMP P1 not fused basally; 12) MP2-IMP similar in length, not fused; 13) foretarsal claws of male not so expanded distally (as in Asthenopodes). Our phylogenetic analyses only recovered some of these character states as synapomorphies of this genus (see diagnosis and Appendix 2). The proposal of fnins.2013.00251 Dom guez (1988) and Hubbard Dom guez (1988) concerning the intermediacy of Priasthenopus gilliesi with respect to Asthenopus curtus and Asthenopodes picteti is in concordance with our benefits. Priasthenopus gilliesi resulted sister for the Povilla-Asthenopus clade, presenting some plesiomorphic character states shared with Asthenopodes. Essential towards the species of Asthenopus Male 1 ?Penile lobe (distad to basal thumb) with a related width along its length, basal thumb separated by a wide furrow (Figs 17A , E ); fnhum.2013.00686 forceps very stout (ratio length/ basal width = 4.7?.0) ...........................................................2 Penile lobe (distad to basal thumb) wider basally, basal thumb fused to penile lobe (Fig. 17D,G); forceps relatively slender (ratio length/ basal width = six.2?.0) ......................................................................................................Phylogeny and biogeography of Asthenopodinae using a revision of Asthenopus...Figure 16. Asthenopus fore (FW) and hind wings (HW) of male imago.