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More detailed experiments will be necessary to further study this finding. As mentioned earlier only MMP23B a small change was noted in CH4 concentration in the headspace above untreated cells during the initial 24?h incubation period (see Figs?2 & 3). It has previously been reported that stable carbon isotope measurements are much more sensitive in detecting CH4 emissions from plant material than by measurement of their concentration (Vigano et?al. 2008; Br��ggemann et?al. 2009). For example, CH4 emission rates as low as 0.03?ng CH4 g?1 DW were measured from 99% 13C-labelled dried wheat (Triticum aestivum) leaves (Vigano et?al. 2008). Thus, we decided to also measure the ��13C value of CH4 in the headspace above 13C isotopically labelled tobacco cells (see Methods section) without any treatment before and after incubation. Figure?4a shows that the ��13C(CH4) values for the headspace above the medium and unlabelled cells did not change over the entire incubation period and closely reflected the ��13C(CH4) of the initial value of laboratory background air (?46.1?��?0.3��). Similar values of ��13C(CH4) in the headspace of the vials (ranging from ?45.9 to ?46.3��) were measured for all samples after incubation for 3?h and these values did not change in control samples (medium only) throughout the rest of the 24?h incubation period. In contrast there was a change in ��13C(CH4) check details values towards higher values (P? same range to those reported (0.16�C0.7?ng?g?1 DW h?1) for grey poplar shoot cultures grown under low light sterile conditions Ibrutinib (Br��ggemann et?al. 2009). In another set of experiments in which the 13C labelled tobacco cell cultures were treated with 2.5?mM NaN3 a strong and highly significant (P?