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Conflict of Interest Statement The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest. Acknowledgments This research was supported by the National Science Foundation Frontiers in Integrative Biology Research Program (EF-0328698), the National Aeronautics and Space Administration Exobiology Program (NNX09AM87G), the Danish Council for Independent Research | Natural Sciences (to MK and SJ), and the U.S. Department of Energy (DOE), Office of Biological and Environmental Research (BER), as part of BER��s Genomic Science Program 395 (GSP). This contribution originates from the GSP RRAD Foundational Scientific Focus Area (FSFA) at the Pacific Northwest National Laboratory (PNNL) under contract 112443. We appreciate support from the Montana Agricultural Experiment Station (project 911352). This study was conducted under Yellowstone National Park research permits YELL-0129 and 5494 (DW) and YELL-02058 (MK), and we appreciate the assistance from National Park Service personnel. We thank Jennifer Weeding from Montana State University for her help with the statistical analysis, and we thank Manolis Kaparakis of Wesleyan University for help with Stata. Footnotes 1http://454.com/downloads/my454/documentation/gs-flx-plus/Rapid-Library-Preparation-Method-Manual_XLPlus_May2011.pdf. 2http://github.com/sandain/pigeon 3http://fcohan.web.wesleyan.edu/ecosim/ Supplementary Material The Supplementary Material for this article can be found online at: http://journal.frontiersin.org/article/10.3389/fmicb.2015.00590 Click here for additional data file.(1.9M, DOC)""Thermococcus kodakarensis KOD1 is a hyperthermophilic anaerobic archaeon, isolated from a solfatara (102��C, pH 5.8) on the shore of Kodakara Island, Kagoshima, Japan (Morikawa et al., 1994). The environmental conditions are not always conducive to steady growth, as fluctuations in temperature regime, fluid flux, and carbon substrate supply create a spatial and temporal mosaic of microenvironments (Edgcomb et al., 2007). The different environmental conditions over time have facilitated the evolution of Archaea for adaptation to extreme environments, and indeed, these bacteria experience difficulties acclimating to less extreme conditions (Reed et al., 2013). T. kodakarensis KOD1 senses the environment and responds to changing environmental conditions (Izumi et al., 2001). Many proteins have been reported to play important roles in cellular protection against different stresses. For example, osmotically inducible protein C (OsmC) from T.